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Diskuzní fórum pro dívky, které zrovna řeší nějaký problém, nebo si jen chtějí popovídat o vztazích, kosmetice, líčení, módě a nákupech, hudbě, filmech nebo o čemkoliv jimém.
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October 2012. Anna Martis Paper on the Retreats is now online. Please find below a link to Annas excellent paper on the Retreat of the Lincoln Washington battalion from Batea to Corbera http://www.albavolunteer.org/2012/07/in-the-footsteps-of-the-lincoln-washington-battalion/ ___________________________________________________________ 31st March and 1st April 2012. 74th Anniversary Talk and Walk concerning the Retreat of the Lincoln Washington Battalion from Batea…
1. Cytochrome alpha 3 in whole-cell suspensions of the fission yeast Schizosaccharomyces pombe reacted in the reduced form with CO to give a photodissociable CO complex with absorption maxima at 429, 543 and 591 nm in CO-liganded reduced-minus-reduced difference spectra. 2. Other CO-bound haemoproteins, cytochromes P-420 and P-450, were not photodissociated under the conditions employed. 3. Measurements of the rates of reassociation of CO with cytochrome alpha 3 after flash photolysis over the temperature range from −101 to −109 degrees C gave a value for Eact. of 28.6 kJ/mol. 4. Between −94 and −106 degrees C, O2 reacted with cytochrome oxidase in intact cells to give an oxygenated intermediate (compound A). 5. At −70 degrees C compound A was converted into a second spectrally distinct intermediate (compound B). 6. Electron transport, indicated by the oxidation of cytochromes alpha + alpha 3 and cytochrome c, did not occur until the temperature was raised to −50 degrees C. 7. At ...
TY - JOUR. T1 - Degradation of HMG-CoA reductase-induced membranes in the fission yeast, Schizosaccharomyces pombe. AU - Lum, Pek Yee. AU - Wright, Robin. PY - 1995/10/1. Y1 - 1995/10/1. N2 - Elevated levels of certain membrane proteins, including the sterol biosynthetic enzyme HMG-CoA reductase, induce proliferation of the endoplasmic reticulum. When the amounts of these proteins return to basal levels, the proliferated membranes are degraded, but the molecular details of this degradation remain unknown. We have examined the degradation of HMG-CoA reductase-induced membranes in the fission yeast, Schizosaccharomyces pombe. In this yeast, increased levels of the Saccharomyces cerevisiae HMG-CoA reductase isozyme encoded by HMG1 induced several types of membranes, including karmellae, which formed a cap of stacked membranes that partially surrounded the nucleus. When expression of HMG1 was repressed, the karmellae detached from the nucleus and formed concentric, multilayered membrane whorls that ...
The fission yeast Schizosaccharomyces pombe divides by medial fission through the use of an actomyosin contractile ring. Precisely at the end of anaphase, the ring begins to constrict and the septum forms. Proper coordination of cell division with mitosis is crucial to ensure proper segregation of chromosomes to daughter cells. The Sid2p kinase is one of several proteins that function as part of a novel signaling pathway required for initiation of medial ring constriction and septation. Here, we show that Sid2p is a component of the spindle pole body at all stages of the cell cycle and localizes transiently to the cell division site during medial ring constriction and septation. A medial ring and an intact microtubule cytoskeleton are required for the localization of Sid2p to the division site. We have established an in vitro assay for measuring Sid2p kinase activity, and found that Sid2p kinase activity peaks during medial ring constriction and septation. Both Sid2p localization to the division site
In this study, we created a fission yeast insertion mutant library in which all mutants were tagged with unique barcode sequences and stored as two readily available selection platforms. The 384-well mutant arrays allow genetic screens on individual mutants and can be extended to genetic approaches such as synthetic genetic array (SGA) [45, 46]. These mutant arrays have been used to identify mutants with four distinct phenotypes (Table 2) as well as strains that are hyper-sensitive to cancer chemotherapeutics camptothecin and bleomycin (Hale and Runge, unpublished data). In addition to 384-well mutant arrays, mutant pools of 1800 mutants are available for parallel analysis.. The insertion mutagenesis used in this study relied on random non-homologous recombination, where a vast majority of transformants have unstable, circularized vector DNA and only a small portion have stable insertions. To facilitate the collection of stable insertion mutants, we included low-dose 5-FOA in our initial ...
Syntaxin is a component of t-soluble N-ethylmaleimide-sensitive factor attachment protein receptor (SNARE), which is responsible for docking membrane vesicles at the target membrane and is highly conserved among eukaryotes. In the fission yeast Schizosaccharomyces pombe, the psy1(+) gene encoding a …
Cytokinesis is a crucial event in cell division. It needs to occur at the right time, in the right place, and only once during a single cell cycle. As such, it must be highly regulated. We have presented evidence showing that par1- and par2-directed PP2A activity is required for proper functioning of the S. pombe SIN pathway. This activity is required at the level of the Ras-like GTPase, Spg1p, whose function is situated at the beginning of this signaling pathway. It appears that the role played by par1 and par2 is to negatively regulate SIN, thereby ensuring that multiple rounds of septation do not occur.. The suppression of spg1-106 by par1Δ is specific to the restoration of the SIN pathway function: As a strain with a loss-of-function ts allele of spg1, spg1-106 cells displayed a typical sid phenotype when shifted to 36°, their restrictive temperature. Deleting par1 in this strain strongly suppressed both the morphological and growth defects of spg1-106 cells. When par1Δ spg1-106 double ...
NUCLEOCYTOPLASMIC transport is a process specific to eukaryotes in which the chromosomes are physically separated from the cytoplasm by the nuclear envelope (NE). In all eukaryotes, the receptor-mediated transport of nuclear proteins across the NE from their site of synthesis in the cytoplasm is essential for all nuclear processes. The precise tissue-specific and temporal regulation of nuclear protein import is also critical in the regulation of cell cycle progression and in developmental and signal transduction pathways (reviewed in Kaffman and OShea 1999).. Proteins are targeted to the nucleus by an NLS (nuclear localization signal). There are two types of classical NLSs, both of which must bind to an importin-α adaptor for transport to the nucleus: the mono-partite NLS that consists of 4 or more basic amino acids preceded by a helix-breaking residue and the bipartite NLS that has two short stretches of basic amino acids separated by a 9- to 12-amino-acid spacer (reviewed in Izaurralde and ...
Elevated levels of certain membrane proteins, including the sterol biosynthetic enzyme HMG-CoA reductase, induce proliferation of the endoplasmic reticulum. When the amounts of these proteins return to basal levels, the proliferated membranes are degraded, but the molecular details of this degradation remain unknown. We have examined the degradation of HMG-CoA reductase-induced membranes in the fission yeast, Schizosaccharomyces pombe. In this yeast, increased levels of the Saccharomyces cerevisiae HMG-CoA reductase isozyme encoded by HMG1 induced several types of membranes, including karmellae, which formed a cap of stacked membranes that partially surrounded the nucleus. When expression of HMG1 was repressed, the karmellae detached from the nucleus and formed concentric, multilayered membrane whorls that were then degraded. During the degradation process, CDCFDA-stained compartments distinct from preexisting vacuoles formed within the interior of the whorls. In addition to these compartments, ...
The fission yeast Schizosaccharomyces pombe is a useful model for analysing DNA replication as genetic methods to allow conditional inactivation of relevant proteins can provide important information about S-phase execution. A number of strategies are available to allow regulation of protein level or activity but there are disadvantages specific to each method and this may have limitations for particular proteins or experiments. We have investigated the utility of the inducible hormone-binding domain (HBD) system, which has been described in other organisms but little used in fission yeast, for the creation of conditional-lethal replication mutants. In this method, proteins are tagged with HBD and can be regulated with β-estradiol. In this article, we describe the application of this method in fission yeast, specifically with regard to analysis of the function of GINS, an essential component of the eukaryotic replicative helicase, the CMG complex.
To decrypt the regulatory code of the genome, sequence elements must be defined that determine the kinetics of RNA metabolism and thus gene expression. Here, we attempt such decryption in an eukaryotic model organism, the fission yeast S. pombe. We first derive an improved genome annotation that redefines borders of 36% of expressed mRNAs and adds 487 non-coding RNAs (ncRNAs). We then combine RNA labeling invivo with mathematical modeling to obtain rates of RNA synthesis and degradation for 5,484 expressed RNAs and splicing rates for4,958 introns. We identify functional sequence elements inDNA and RNA that control RNA metabolic rates and quantifythecontributions of individual nucleotides to RNA synthesis,splicing, and degradation. Our approach reveals distinct kineticsof mRNA and ncRNA metabolism, separates antisense regulation by transcription interference from RNA interference, and provides a general tool for studying the regulatory code of genomes. ...
The cdc2+ gene function plays a central role in the control of the mitotic cell cycle of the fission yeast Schizosaccharomyces pombe. Recessive temperature-sensitive mutations in the cdc2 gene cause cell cycle arrest when shifted to the restrictive temperature, while a second class of mutations within the cdc2 gene causes a premature advancement into mitosis. Previously the cdc2+ gene has been cloned and has been shown to encode a 34 kDa phosphoprotein with in vitro protein kinase activity. Here we describe the cloning of 11 mutant alleles of the cdc2 gene using two simple methods, one of which is presented here for the first time. We have sequenced these alleles and find a variety of single amino acid substitutions mapping throughout the cdc2 protein. Analysis of these mutations has identified a number of regions within the cdc2 protein that are important for cdc2+ activity and regulation. These include regions which may be involved in the interaction of the cdc2+ gene product with the proteins ...
Inactivating a specific protein in vivo can yield important information about its function. One strategy previously developed in Saccharomyces cerevisiae by the Varshavsky group involves fusing a degron, derived from mouse dihydrofolate reductase, to the N-terminus of the target protein, which thereby confers temperature-sensitive degradation at the restrictive temperature. We describe here the application of this technique in the fission yeast, Schizosaccharomyces pombe.
Eukaryotic cells have developed elaborate regulatory mechanisms to ensure that DNA replication is restricted to S phase and occurs just once per cell cycle. An important component of this regulation is the coordination of multiple origins of replication. Data from many systems have provided a model of the initiation of DNA synthesis in which several multiprotein complexes interact with discrete replication origins in a specific temporal pattern to regulate entry into S phase (24, 26, 32). Two protein kinase complexes are required to activate the replication origins: cyclin-cyclin-dependent kinase (CDK), which acts both positively and negatively to control origin function (21,22), and the product of the hsk1 + (also called CDC7) gene complex.. Schizosaccharomyces pombe hsk1 + is a member of the conserved family of CDC7 protein kinases (23, 37). Thehsk1 + gene was originally cloned by its sequence homology to budding yeast CDC7 and is essential for viability (36). Spores with hsk1 +deleted are ...
Eukaryotic cells have developed elaborate regulatory mechanisms to ensure that DNA replication is restricted to S phase and occurs just once per cell cycle. An important component of this regulation is the coordination of multiple origins of replication. Data from many systems have provided a model of the initiation of DNA synthesis in which several multiprotein complexes interact with discrete replication origins in a specific temporal pattern to regulate entry into S phase (24, 26, 32). Two protein kinase complexes are required to activate the replication origins: cyclin-cyclin-dependent kinase (CDK), which acts both positively and negatively to control origin function (21,22), and the product of the hsk1 + (also called CDC7) gene complex.. Schizosaccharomyces pombe hsk1 + is a member of the conserved family of CDC7 protein kinases (23, 37). Thehsk1 + gene was originally cloned by its sequence homology to budding yeast CDC7 and is essential for viability (36). Spores with hsk1 +deleted are ...
In response to diverse environmental stresses, the phosphorylation of eukaryotic initiation factor-2 (eIF2α) induces a programme of gene expression that mitigates cellular injury. Several protein kinases phosphorylate eIF2α at serine 51, leading to the inhibition of eIF2B activity, that in turn causes a reduction in general protein synthesis and an enhancement of the translation of specific mRNAs encoding for proteins that remediate the stress. In mammalian cells, four eIF2 kinases (HRI, GCN2, PEK/Perk and PKR) inhibit translation initiation through the phosphorylation of eIF2α in response to different types of cellular stress (Dever, 2002; Proud, 2005). In Saccharomyces cerevisiae, Gcn2p is activated upon nutrient limitation (amino acids, purine and glucose), but also by high concentrations of sodium, rapamycin and methyl methanesulfonate (Cherkasova and Hinnebusch, 2003; Narasimhan et al., 2004; Valenzuela et al., 2001; Yang et al., 2000). In the fission yeast Schizosaccharomyces pombe, ...
In virtually all eukaryotes, mitosis starts after the completion of DNA synthesis. This orderly process is ensured by the checkpoint mechanism that blocks the onset of mitosis while DNA is being synthesized or is damaged. In the fission yeast Schizosaccharomyces pombe, this mechanism involves some r …
Reactive oxygen species (ROS) can damage cellular components leading to cell death, but paradoxically, ROS also play essential roles in metabolism and signalling in eukaryotic cells. Dysregulation of this balance is associated with a range of host diseases and cells have consequently evolved sophisticated signalling networks to sense, detoxify and adapt to changes in ROS levels. Hydrogen peroxide, for example, is reduced by thiol-peroxidases which in turn, can trigger the oxidation of thiol-dependent redox transcription factors. However, the relationship between hydrogen peroxide stimuli and the level of redox transcription factor activation has largely been described in qualitative terms. Because quantitative measures of the redox signal have been lacking, we tested whether three signalling parameters viz. the signalling time, duration and amplitude could be used to quantify the hydrogen peroxide-dependent redox signal in the Tpx1/Pap1 pathway in Schizosaccharomyces pombe. We found significant ...
Kohli, J., Hottinger, H., Munz, P., Strauss, A., Thuriaux, P. 1977. Genetic mapping in Schizosaccharomyces pombe by mitotic and meiotic analysis and induced haploidization. Genetics, 87, 471-489 ...
The Schizosaccharomyces pombe HIRA-like protein hip1 is required for the periodic expression of histone genes and contributes to the function of complex ...
Brown, William R.A. and Litti, Gianni and Rosa, Carlos and James, Steve and Roberts, Ian and Robert, Vincent and Jolly, Neil and Tang, Wen and Baumann, Peter and Green, Carter and Schlegel, Kristina and Young, Jonathan and Hirchaud, Fabienne and Leek, Spencer and Thomas, Geraint and Blomberg, Anders and Warringer, Jonas (2011) A geographically diverse collection of schizosaccharomyces pombe isolates shows limited phenotypic variation but extensive karyotypic diversity. G3, 1 (7). pp. 615-626. ISSN 2160-1836 ...
TY - JOUR. T1 - Cell-cycle control linked to extracellular environment by MAP kinase pathway in fission yeast. AU - Shiozaki, Kazuhiro. AU - Russell, Paul. PY - 1995/12/14. Y1 - 1995/12/14. N2 - In fission yeast the onset of mitosis is brought about by Cdc2/Cdc13 kinase, which is inhibited by the Wee1/Mik1 tyrosine kinases and activated by Cdc25 tyrosine phosphatase. This control network integrates many signals, including those that monitor DNA replication, DNA damage and cell size. We report here that a fission yeast MAP kinase pathway links the cell-cycle G2/M control with changes in the extracellular environment that affect cell physiology. Fission yeast spc1- mutants have a G2 delay that is greatly exacerbated by growth in high osmolarity media and nutrient limitation. A lethal interaction of spc1 and cdc25 mutations shows that Spc1 promotes the onset of mitosis. Spc1 is a MAP kinase homologue that is activated by Wis1 kinase in response to osmotic stress and nutrient limitation. Spc1 is ...
Binds specifically to cytosolic chaperonin (c-CPN) and transfers target proteins to it. Binds to nascent polypeptide chain and promotes folding in an environment in which there are many competing pathways for nonnative proteins (By similarity).
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Project Information The Schizosaccharomyces Comparative Genome Project project is part of the Broad Institute Fungal Genome Initiative. Its goal was to sequence, annotate and analyze the genomes and transcriptomes of Schizosaccharomyces japonicus, Schizosaccharomyces octosporus and Schizosaccharomyces cryophilus. The primary collaborator for this project was Nick Rhind at the University of Massachusetts, Worcester, Medical School.
ID AB009602; SV 1; linear; mRNA; STD; FUN; 561 BP. XX AC AB009602; XX DT 15-DEC-1997 (Rel. 53, Created) DT 14-APR-2005 (Rel. 83, Last updated, Version 2) XX DE Schizosaccharomyces pombe mRNA for MET1 homolog, partial cds. XX KW MET1 homolog. XX OS Schizosaccharomyces pombe (fission yeast) OC Eukaryota; Fungi; Dikarya; Ascomycota; Taphrinomycotina; OC Schizosaccharomycetes; Schizosaccharomycetales; Schizosaccharomycetaceae; OC Schizosaccharomyces. XX RN [1] RP 1-561 RA Kawamukai M.; RT ; RL Submitted (07-DEC-1997) to the EMBL/GenBank/DDBJ databases. RL Makoto Kawamukai, Shimane University, Life and Environmental Science; 1060 RL Nishikawatsu, Matsue, Shimane 690, Japan RL (E-mail:[email protected], Tel:0852-32-6587, Fax:0852-32-6499) XX RN [2] RP 1-561 RA Kawamukai M.; RT "S.pmbe MET1 homolog"; RL Unpublished. XX FH Key Location/Qualifiers FH FT source 1..561 FT /organism="Schizosaccharomyces pombe" FT /mol_type="mRNA" FT /clone_lib="pGAD GH" FT /db_xref="taxon:4896" FT CDS ,1..275 FT ...
Dr. Nurses research focuses on the molecular machineries that control cell division and cell shape. Using the fission yeast Schizosaccharomyces pombe as a model system, his laboratory studies the cell cycle and cell morphogenesis controls operative in eukaryotic cells. His major past contribution was the codiscovery of cyclin-dependent kinase (CDK) as the key regulator molecule controlling S phase and mitosis, findings that have had implications for understanding cell reproduction, cell growth, development, and cancer. Present work in the Nurse laboratory is in three areas: the cell cycle, cell form, and genomic studies. The lab is split on two sites, with the major activity located at the Francis Crick Institute in London, and a smaller group located at The Rockefeller University, which works mainly on combining chemical biology and genetics to investigate cell biology problems in fission yeast.. In collaboration with Tarun Kapoor, the Nurse lab works on the development and use of fission ...
Progression through the eukaryotic cell cycle is governed by cyclin-dependent kinases (CDKs) in conjunction with their cyclin partners (Morgan, 1997). (For nomenclature, Arabidopsis thaliana wild-type genes [in italics] and proteins [not italicized] use all uppercase letters, and mutants are in lowercase letters [in italics]. In fission yeast [Schizosaccharomyces pombe] and budding yeast [Saccharomyces cerevisiae], wild-type proteins are not italicized and in title case, with fission yeast having a postfix superscript "+" and budding yeast a postfix lowercase "p" indicating the protein nature. All other proteins are in title case except if stated otherwise in the National Center for Biotechnology Information [NCBI] library. Generalized terms appear not italicized and uppercase.) A key principle and the driving force of the cell cycle is the generation of alternating phases of high and low kinase activity (Nasmyth, 1996). At times of high CDK-cyclin activity, a number of target proteins become ...
Bioneers exclusive Schizosaccharomyces pombe (S. pombe) Genome-wide Deletion Mutant Library is a powerful tool for large-scale genetic functional analysis, identification and verification research of drug targets and for integrated systems research of cell function. Co-developed by Bioneer and KRIBB in collaboration with Dr. Paul Nurse of the Cancer Research Center in UK, the S. pombe Genome-wide Deletion Mutant Library can be used for genetic and chemical screening such as drug target identification, gene expression profiling, and synthetic lethal profiling. S. pombe offers higher homology with mammalian cells and human genes than those of S. pombe. S. pombe Genome-wide Deletion Mutant Library targets every ORF (4,914 types) in the S. pombe genome through a targeted mutagenesis method. A total of 4,836 heterozygous diploid deletion mutants representing 98.4% of the organism genome and 3,400 haploid deletion mutants with 95.3% genome coverage are available. Since there are different tag ...
Win, Thein Zaw; (2001) Characterisation of two Class V myosins in the fission yeast Schizosaccharomyces pombe. Doctoral thesis (Ph.D), UCL (University College London). Green open ...
Eukaryotic organisms use cell-cycle checkpoints to ensure that nuclear division is restrained while DNA is undergoing replication or repair. Recent studies of the fission yeast Schizosaccharomyces pombe have illuminated these checkpoint mechanisms. These investigations have connected checkpoint proteins with central elements of the mitotic-control machinery ...
Protein kinase that acts both on serines and on tyrosines. It acts as a dosage-dependent negative regulator of entry into mitosis (G2 to M transition). Phosphorylates and inhibits cdc2.
Fission yeast cell division is initiated by the cdc2/cdc13-cyclin protein kinase which in its catalytically active state comprises the mitotic inducer. During interphase the cdc2/cyclin complex is assembled in an inactive state that requires cdc25+ gene function for M-phase activation. The cdc25+ product, a 76 kd phosphoprotein, is shown to oscillate in abundance during the cell cycle, reaching a peak at G2/M, and to be sensitive to nitrogen starvation. The level of cdc25 is subject to feedback regulation involving both cdc25 and cdc2.. ...
We are interested in understanding how gene expression is encoded in genomes, and how to leverage this knowledge to understand the basis of genetic diseases. To this end, we employ statistical modeling of omics data and work in close collaboration with experimentalists. I will provide an overview of recent studies from our lab, covering work on RNA metabolism in S. pombe [1] and on mRNA stability in S. cerevisiae [2], and discussing the implications of TT-seq, a novel protocol suited to study nascent transcriptomes in higher eukaryotes [3]. I will also report on a recent pilot study where we established RNA-sequencing as a powerful companion tool to genome sequencing for pinpointing molecular causes of rare genetic disorders [4]. The talk will end with a discussion on future directions in computational modeling of cis-regulatory elements.. References [1] Eser. et al, Determinants of RNA metabolism in the Schizosaccharomyces pombe genome, Molecular Systems Biology, 2016 [2] Cheng, et al. ...
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Hello I would like to get some information considering the growth of S. pombe. It is known that S. pombe cells divide by medial fission. In effect each of the mother cell splits into two identical doughter cells. On the other hand there are some information about possibility of age asymetry in the case of S. pombe cell fission. So can we say that S. pombe divides asymetrically (at least in some conditions)? Can we make a distinction between mother and dougter cell in case of S. pombe ? I would be grateful for any suggestions. Thanks Magdalena Frajnt ...
A comprehensive formalism to calculate fission cross sections based on the extension of the optical model for fission is presented. It can be used for description of nuclear reactions on actinides featuring multi-humped fission barriers with partial absorption in the wells and direct transmission through discrete and continuum fission channels. The formalism describes the gross fluctuations observed in the fission probability due to vibrational resonances, and can be easily implemented in existing statistical reaction model codes. The extended optical model for fission is applied for neutron induced fission cross-section calculations on 234,235,238U and 239Pu targets. A triple-humped fission barrier is used for 234,235U(n,f), while a double-humped fission barrier is used for 238U(n,f) and 239Pu(n,f) reactions as predicted by theoretical barrier calculations. The impact of partial damping of class-II/III states, and of direct transmission through discrete and continuum fission channels, is shown ...
Cell, Cell Division, Cell Proliferation, Cells, Fission Yeast, Germ Cell, Heterochromatin, Kinases, Oncogenesis, Regulation, Schizosaccharomyces, Schizosaccharomyces Pombe, Yeast
S pombe RST2 protein: regulates the ste11 gene; isolated from Schizosaccharomyces pombe; homologous to S. cerevisiae ADR1 protein; amino acid sequence in first source
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Modules of approx. 60 residues attached to a few GH81 enzymes but present in other proteins. Binding to β-1,3-glucan demonstrated for Schizosaccharomyces pombe 972h- endo-1,3-β-glucanase Eng1 [Martin-Quadrado et al., Mol. Microbiol. (2008) 69:188-200 PMID:18466295 ...